The Life of the Fly

第9章

CHAPTERXTHEGREYFLESHFLIES

Herethecostumechanges,notthemanneroflife。Wefindthesamefrequentingofdeadbodies,thesamecapacityforthespeedyliquefactionofthefleshymatter。Iamspeakingofanash—grayfly,thegreenbottle’ssuperiorinsize,withbrownstreaksonherbackandsilvergleamsonherabdomen。Notealsotheblood—redeyes,withthehardlookoftheknackerinthem。ThelanguageofscienceknowsherasSarcophaga,theflesheater;inthevulgartonguesheisthegreyfleshfly,orsimplythefleshfly。

Letnottheseexpressions,howeveraccurate,misleadusintobelievingforamomentthattheSarcophagaearetheboldcompanyofmastertainterswhohauntourdwellings,moreparticularlyinautumn,andplanttheirvermininourill—guardedviands。TheauthorofthoseoffencesisCalliphoravomitoria,thebluebottle,whoisofastouterbuildandarrayedindarkestblue。Itisshewhobuzzesagainstourwindowpanes,whocraftilybesiegesthemeatsafeandwholiesinwaitinthedarknessforanopportunitytooutwitourvigilance。Theother,thegreyfly,worksjointlywiththegreenbottles,whodonotventureinsideourhousesandwhoworkinthesunlight。Lesstimid,however,thanthey,shouldtheoutdooryieldbesmall,shewillsometimescomeindoorstoperpetratehervillainies。Whenherbusinessisdone,shemakesoffasfastasshecan,forshedoesnotfeelathomewithus。

Atthismoment,mystudy,averymodestextensionofmyopenairestablishments,hasbecomesomethingofacharnelhouse。Thegreyflypaysmeavisit。IfIlayapieceofbutcher’smeatonthewindowsill,shehastensup,worksherwillonitandretires。Nohidingplaceescapeshernoticeamongthejars,cups,glassesandreceptaclesofeverykindwithwhichmyshelvesarecrowded。

Withaviewtocertainexperiments,Icollectedaheapofwaspgrubs,asphyxiatedintheirundergroundnests。Stealthilyshearrives,discoversthefatpileand,hailingastreasuretrovethisprovenderwhereofherraceperhapshasnevermadeusebefore,entruststoitaninstallmentofherfamily。Ihaveleftatthebottomofaglassthebestpartofahard—boiledeggfromwhichI

havetakenafewbitsofwhiteintendedforthegreenbottlemaggots。Thegreyflytakespossessionoftheremains,recksnotoftheirnoveltyandcolonizesthem。Everythingsuitsherthatfallswithinthecategoryofalbuminousmatters:everything,downtodeadsilkworms;everything,downtoamessofkidney—beansandchick—peas。

Nevertheless,herpreferenceisforthecorpse:furredbeastandfeatheredbeast,reptileandfish,indifferently。Togetherwiththegreenbottles,sheissedulousinherattendanceonmypans。

Dailyshevisitsmysnakes,takesnoteoftheconditionofeachofthem,savorsthemwithherproboscis,goesaway,comesback,takeshertimeandatlastproceedstobusiness。Still,itisnothere,amidthetumultofcallers,thatIproposetofollowheroperations。Alumpofbutcher’smeatlaidonthewindowsill,infrontofmywritingtable,willbelessoffensivetotheeyeandwillfacilitatemyobservations。

TwofliesofthegenusSarcophagafrequentmyslaughteryard:

SarcophagacarnariaandSarcophagahaemorrhoidalis,whoseabdomenendsinaredspeck。Thefirstspecies,whichisalittlelargerthanthesecond,ismorenumerousanddoesthebestpartoftheworkintheopenairshamblesofthepans。Itisthisflyalsowho,atintervalsandnearlyalwaysalone,hastenstothebaitexposedonthewindowsill。

Shecomesupsuddenly,timidly。SoonshecalmsherselfandnolongerthinksoffleeingwhenIdrawnear,forthedishsuitsher。

Sheissurprisinglyquickaboutherwork。Twiceover——buzz!Buzz!—

—thetipofherabdomentouchesthemeat;andthethingisdone:agroupofverminwrigglesout,releasesitselfanddispersessonimblythatIhavenotimetotakemylensandcountthenaccurately。Asseenbythenakedeye,therewereadozenofthem。

Whathasbecomeofthem?Onewouldthinkthattheyhadgoneintotheflesh,attheveryspotwheretheywerelaid,soquicklyhavetheydisappeared。Butthatdiveintoasubstanceofsomeconsistencyisimpossibletothesenewbornweaklings。Wherearethey?Ifindthemmoreorlesseverywhereinthecreasesofthemeat;singlyandalreadygropingwiththeirmouths。Tocollecttheminordertonumberthemisnotpracticable,forIdonotwanttodamagethem。Letusbesatisfiedwiththeestimatemadeatarapidglance:thereareadozenorso,broughtintotheworldinonedischargeofalmostinappreciablelength。

Thoselivegrubs,takingtheplaceoftheusualeggs,havelongbeenknown。Everybodyisawarethatthefleshfliesbringforthlivingmaggots,insteadoflayingeggs。Theyhavesomuchtodoandtheirworkissourgent!Tothem,theinstrumentsofthetransformationofdeadmatter,adaymeansaday,alongspaceoftimewhichitisallimportanttoutilize。Thegreenbottle’seggs,thoughtheseareofveryrapiddevelopment,taketwenty—fourhourstoyieldtheirgrubs。Thefleshfliessaveallthistime。Fromtheirmatrix,laborersflowstraightwayandsettoworkthemomenttheyareborn。Withtheseardentpioneersofsanitation,thereisnorestattendantuponthehatching,thereisnotaminutelost。

Thegang,itistrue,isnotanumerousone;buthowoftencanitnotberenewed!ReadReaumur’sdescriptionofthewonderfulprocreatingmachineryboastedbytheFleshflies。Itisaspiralribbon,avelvetyscrollwhosenapisasortoffleeceofmaggotssetcloselytogetherandeachcasedinasheath。Thepatientbiographercountedthehost:itnumbers,hetellsus,nearlytwentythousand。Youareseizedwithstupefactionatthisanatomicalfact。

Howdoesthegrayflyfindthetimetosettleafamilyofsuchdimensions,especiallyinsmallpackets,asshehasjustdoneonmywindowsill?Whatanumberofdeaddogs,molesandsnakesmustshenotvisitbeforeexhaustingherwomb!Willshefindthem?Corpsesofmuchsizedonotaboundtothatextentinthecountry。Aseverythingsuitsher,shewillalightonotherremainsofminorimportance。Shouldtheprizebearichone,shewillreturntoittomorrow,thedayafterandlaterstill,overandoveragain。Inthecourseoftheseason,bydintofpacketsofgrubsdepositedhere,thereandeverywhere,shewillperhapsendbyhousingherentirebrood。Butthen,ifallthingsprosper,whataglut,forthereareseveralfamiliesbornduringtheyear!Wefeelitinstinctively:theremustbeachecktothesegenerativeenormities。

Letusfirstconsiderthegrub。Itisasturdymaggot,easytodistinguishfromthegreenbottle’sbyitslargergirthandespeciallybythewayinwhichitsbodyterminatesbehind。Thereishereasuddenbreakingoff,hollowedintoadeepcup。Atthebottomofthiscrateraretwobreathingholes,twostigmatawithamber—redtips。Theedgeofthecavityisfringedwithhalfascoreofpointed,fleshyfestoons,whichdivergelikethespikesofacoronet。Thecreaturecancloseoropenthisdiadematwillbybringingthedenticulationstogetherorbyspreadingthemoutwide。

Thisprotectstheairholeswhichmightotherwisebechokedupwhenthemaggotdisappearsintheseaofbroth。Asphyxiawouldsupervene,ifthetwobreathingholesatthebackbecameobstructed。Duringtheimmersion,thefestoonedcoronetshutslikeaflowerclosingitspetalsandtheliquidisnotadmittedtothecavity。

Nextfollowstheemergence。Thehindpartreappearsintheair,butappearsalone,justatthelevelofthefluid。Thenthecoronetspreadsoutafresh,thecupgapesandassumestheaspectofatinyflower,withthewhitedenticulationsforpetalsandthetwobrightreddots,thestigmataatthebottom,forstamens。Whenthegrubs,pressedoneagainsttheother,withtheirheadsdownwardsinthefetidsoup,makeanunbrokenshoal,thesightofthosebreathingcupsincessantlyopeningandclosing,withalittleclacklikeavalve,almostmakesoneforgetthehorrorsofthecharnelyard。ItsuggestsacarpetoftinySeaanemones。Themaggothasitsbeautiesafterall。

Itisobvious,iftherebeanylogicinthings,thatagrubsowell—protectedagainstasphyxiationbydrowningmustfrequentliquidsurroundings。Onedoesnotencircleone’shindquarterswithacoronetforthesolesatisfactionofdisplayingit。Withitsapparatusofspokes,theGreyFly’sgrubinformsusofthedangerousnatureofitsfunctions:whenworkinguponacorpse,itrunstheriskofdrowning。Howisthat?Rememberthegrubsofthegreenbottle,fedonhard—boiledwhiteofegg。Thedishsuitsthem;

only,bytheactionoftheirpepsin,itbecomessofluidthattheydiesubmerged。Becauseoftheirhinderstigmata,whichareactuallyontheskinanddevoidofanydefensivemachinery,theyperishwhentheyfindnosupportapartfromtheliquid。

Thefleshfly’smaggots,thoughincomparableliquefiers,knownothingofthisperil,eveninapuddleofcarrionbroth。Theirbulkyhindpartservesasafloatandkeepstheairholesabovethesurface。When,forfurtherinvestigation,theymustneedsgoundercompletely,theanemoneatthebackshutsandprotectsthestigmata。Thegrubsofthegrayflyareendowedwithalifebuoybecausetheyarefirstclassliquefiers,readytoincurthedangerofaduckingatanymoment。

WhenhighanddryonthesheetofcardboardwhereIplacethemtoobservethematmyease,theymoveaboutactively,withtheirbreathingrosewidespreadandtheirstigmatarisingandfallingasasupport。Thecardboardisonmytable,atthreestepsfromanopenwindow,andlitatthistimeofdayonlybythesoftlightofthesky。Well,themaggots,oneandallofthem,turnintheoppositedirectiontothewindow;theyhastily,madlytaketoflight。

Iturnthecardboardround,withouttouchingtherunaways。Thisactionmakesthecreaturesfacethelightagain。Forthwith,thetroopstops,hesitates,takesahalfturnandoncemoreretreatstowardsthedarkness。Beforetheendoftheracecourseisreached,Iagainturnthecardboard。Forthesecondtime,themaggotsveerroundandretracetheirsteps。RepeattheexperimentasoftenasI

will,eachtimethesquadwheelsaboutintheoppositedirectiontothewindowandpersistsinavoidingthetrapoftherevolvingcardboard。

Thetrackisonlyashortone:thecardboardmeasuresthreehand’sbreadthsinlength。Letusgivemorespace。Isettlethegrubsontheflooroftheroom;withahairpencil,Iturnthemwiththeirheadspointingtowardsthelightedaperture。Themomenttheyarefree,theyturnandrunfromthelight。Withallthespeedwhereoftheircripple’sshuffleallows,theycoverthetiledfloorofthestudyandgoandknocktheirheadsagainstthewall,twelvefeetoff,skirtingitafterwards,sometotherightandsometotheleft。Theyneverfeelfarenoughawayfromthathatefulilluminatedopening。

Whattheyareescapingfromisevidentlythelight,for,ifImakeitdarkwithascreen,thetroopdoesnotchangeitsdirectionwhenIturnthecardboard。Itthenprogressesquitereadilytowardsthewindow;but,whenIremovethescreen,itturnstailatonce。

Thatagrubdestinedtoliveinthedarkness,undertheshelterofacorpse,shouldavoidthelightisonlynatural;thestrangepartisitsveryperception。Themaggotisblind。Itspointedforepart,whichwehesitatetocallahead,bearsabsolutelynotraceofanyopticalapparatus;andthesamewitheveryotherpartofthebody。Thereisnothingbutonebare,smooth,whiteskin。Andthissightlesscreature,deprivedofanyspecialnervouspointsservedbyocularpower,isextremelysensitivetothelight。Itswholeskinisasortofretina,incapableofseeing,ofcourse,butable,atanyrate,todistinguishbetweenlightanddarkness。Underthedirectraysofasearchingsun,thegrub’sdistresscouldbeeasilyexplained。Weourselves;withourcoarseskin,incomparisonwiththatofthemaggot,candistinguishbetweensunshineandshadowwithoutthehelpoftheeyes。But,inthepresentcase,theproblembecomessingularlycomplicated。Thesubjectsofmyexperimentreceiveonlythediffusedlightofthesky,enteringmystudythroughanopenwindow;yetthistemperedlightfrightensthemoutoftheirsenses。Theyfleethepainfulapparition;theyarebentuponescapingatallcosts。

Nowwhatdothefugitivesfeel?Aretheyphysicallyhurtbythechemicalradiations?Aretheyexasperatedbyotherradiations,knownorunknown?Lightstillkeepsmanyasecrethiddenfromusandperhapsouropticalscience,bystudyingthemaggot,mightbecomethericherbysomevaluableinformation。Iwouldgladlyhavegonefartherintothequestion,hadIpossessedthenecessaryapparatus。ButIhavenot,IneverhavehadandofcourseInevershallhavetheresourceswhicharesousefultotheseeker。Thesearereservedforthecleverpeoplewhocaremoreforlucrativepoststhanforfairtruths。Letuscontinue,however,withinthemeasurewhichthepovertyofmymeanspermits。

Whendulyfattened,thegrubsofthefleshfliesgoundergroundtotransformthemselvesintopupae。Theburialisintended,obviously,togivethewormthetranquillitynecessaryforthemetamorphosis。Letusaddthatanotherobjectofthedescentistoavoidtheimportunitiesofthelight。Themaggotisolatesitselftothebestofitspowerandwithdrawsfromthegarishdaybeforecontractingintoalittlekeg。Inordinaryconditions,withaloosesoil,itgoeshardlylowerthanahand’sbreadthdown,forprovisionhastobemadeforthedifficultiesofthereturntothesurfacewhentheinsect,nowfullgrown,isimpededbyitsdelicateflywings。Thegrub,therefore,deemsitselfsuitablyisolatedatamoderatedepth。Sideways,thelayerthatshieldsitfromthelightisofindefinitethickness;upwards,itmeasuresaboutfourinches。Behindthisscreenreignsutterdarkness,theburiedone’sdelight。Thisiscapital。

Whatwouldhappenif,byanartifice,thesidewardlayerwerenowherethickenoughtosatisfythegrub?Now,thistime,Ihavethewherewithaltosolvetheproblem,intheshapeofabigglasstube,openatbothends,aboutthreefeetlongandlessthananinchwide。IuseittoblowtheflameofhydrogeninthelittlechemistrylessonswhichIgivemychildren。

Icloseoneendwithacorkandfillthetubewithfine,dry,siftedsand。Onthesurfaceofthislongcolumn,suspendedperpendicularlyinacornerofmystudy,IinstallsometwentySarcophagagrubs,feedingthemwithmeat。Asimilarpreparationisrepeatedinawiderjar,withamouthasbroadasone’shand。Whentheyarebigenough,thegrubsineitherapparatuswillgodowntothedepththatsuitsthem。Thereisnomoretobedonebuttoleavethemtotheirowndevices。

Thewormsatlastburythemselvesandhardenintopupae。Thisisthemomenttoconsultthetwoapparatus。ThejargivesmetheanswerwhichIshouldhaveobtainedintheopenfields。Fourinchesdown,orthereabouts,thewormshavefoundaquietlodging,protectedabovebythelayerthroughwhichtheyhavepassedandoneverysidebythethicknessofthevessel’scontents。Satisfiedwiththesite,theyhavestoppedthere。

Itisaverydifferentmatterinthetube。Theleastburiedofthepupaearehalfayarddown。Othersarelowerstill;mostofthemevenhavereachedthebottomofthetubeandaretouchingthecorkstopper,aninsuperablebarrier。Theselast,wecansee,wouldhavegoneyetdeeperiftheapparatushadallowedthem。Notoneofthescoreofgrubshassettledatthecustomaryhaltingplace;allhavetraveledfartherdownthecolumn,untiltheirstrengthgaveway。Intheiranxiousflight,theyhavedugdeeperandeverdeeper。

Whatweretheyflyingfrom?Thelight。Abovethem,thecolumntraversedformsamorethansufficientshelter;but,atthesides,theirksomesensationisstillfeltthroughacoatofearthhalfaninchthickifthedescentismadeperpendicularly。Toescapethedisturbingimpression,thegrubthereforegoesdeeperanddeeper,hopingtoobtainlowerdowntherestwhichisdenieditabove。Itonlyceasestomovewhenwornoutwiththeeffortorstoppedbyanobstacle。

Now,inasoftdiffusedlight,whatcanbetheradiationscapableofactinguponthisloverofdarkness?Theyarecertainlynotthesimpleluminousrays,forascreenoffine,heapedupearth,nearlyhalfaninchinthickness,isperfectlyopaque。Then,toalarmthegrub,towarnitoftheoverproximityoftheexteriorandsendittomaddepthsinsearchofisolation,otherradiations,knownorunknown,mustberequired,radiationscapableofpenetratingascreenagainstwhichordinaryradiationsarepowerless。Whoknowswhatvistasthenaturalphilosophyofthemaggotmightopenouttous?Forlackofapparatus,Iconfinemyselftosuspicions。

Togoundergroundtoayard’sdepth——andfartherifmytubehadallowedit——isonthepartoftheFleshfly’sgrubavagaryprovokedbyunkindexperiment:neverwoulditburyitselfsolowdown,iflefttoitsownwisdom。Ahand’sbreadththicknessisquiteenough,isevenagreatdealwhen,aftercompletingthetransformation,ithastoclimbbacktothesurface,alaboriousoperationabsolutelyresemblingthetaskofanentombedwellsinker。Itwillhavetofightagainstthesandthatslipsandgraduallyfillsupthesmallamountofemptyspaceobtained;itwillperhaps,withoutcrowbarorpickaxe,havetocutitselfagallerythroughsomethingtantamounttotufa,thatistosay,throughearthwhichashowerhasrenderedcompact。Forthedescent,thegrubhasitsfangs;fortheassent,theflyhasnothing。Onlythatmomentcomeintoexistence,sheisaweakling,withtissuesstilldevoidofanyfirmness。Howdoesshemanagetogetout?Weshallknowbywatchingafewpupaeplacedatthebottomofatest—tubefilledwithearth。ThemethodoftheFleshflieswillteachusthatofthegreenbottlesandtheotherFlies,allofwhommakeuseofthesamemeans。

Enclosedinherpupa,thenascentflybeginsbyburstingthelidofhercasketwithaherniawhichcomesbetweenhertwoeyesanddoublesortreblesthesizeofherhead。Thiscephalicblisterthrobs:itswellsandsubsidesbyturns,owingtothealternatefluxandrefluxoftheblood。Itislikethepistonofanhydraulicpressopeningandforcingbackthefrontpartofthekeg。

Theheadmakesitsappearance。Thehydrocephalousmonstercontinuestheplayofherforehead,whileherselfremainingstationary。Insidethepupa,adelicateworkisbeingperformed:

thecastingofthewhitenymphaltunic。Allthroughthisoperation,theherniaisstillprojecting。Theheadisnottheheadofafly,butaqueer,enormousmitre,spreadingatthebaseintotworedskullcaps,whicharetheeyes。Tosplithercraniuminthemiddle,shuntthetwohalvestotherightandleftandsendsurgingthroughthegapatumorwhichstavesthebarrelwithitspressure:thisconstitutestheFly’seccentricmethod。

Forwhatreasondoesthehernia,oncethekegisstaved,continueswollenandprojecting?Itakeittobeawastepocketintowhichtheinsectmomentarilyforcesbackitsreservesofbloodinordertodiminishthebulkofthebodytothatextentandtoextractitmoreeasilyfromthenymphalsloughandafterwardsfromthenarrowchanneloftheshell。Aslongastheoperationofthereleaselasts,itpushesoutsideallthatitisabletoinjectofitsaccumulatedhumors;itmakesitselfsmallinsidethepupaandswellsintoabloateddeformitywithout。Twohoursandmorearespentinthislaboriousstripping。

Atlast,theflycomesintoview。Thewings,merescantystumps,hardlyreachthemiddleoftheabdomen。Ontheouteredge,theyhaveadeepnotchsimilartothewaistofaviolin。Thisdiminishesbyjustsomuchthesurfaceandthelength,anexcellentdevicefordecreasingthefrictionalongtheearthycolumnwhichhasnexttobescaled。Thehydrocephalousoneresumesherperformancemorevigorouslythanever;sheinflatesanddeflatesherfrontalknob。Thepoundedsandrustlesdowntheinsect’ssides。Thelegsplaybutasecondarypart。Stretchedbehind,motionless,whenthepistonstrokeisdelivered,theyfurnishasupport。Asthesanddescends,theypileitandnimblypushitback,afterwhichtheydragalonglifelesslyuntilthenextavalanche。Theheadadvanceseachtimebyalengthequaltothatofthesanddisplaced。Eachstrokeofthefrontalswellingmeansastepforward。Inadry,loosesoil,thingsgoprettyfast。A

columnsixincheshighistraversedinlessthanaquarterofanhour。

Assoonasitreachesthesurface,theinsect,coveredwithdust,proceedstomakeitstoilet。Itthrustsouttheblisterofitsforeheadforthelasttimeandbrushesitcarefullywithitsfronttarsi。Itisimportantthatthelittlepoundingengineshouldbecarefullydustedbeforeitistakeninsidetoformaforeheadthatwillopennomore:thislestanygritshouldlodgeinthehead。

Thewingsarecarefullybrushedandpolished;theylosetheircurvednotches;theylengthenandspread。Then,motionlessonthesurfaceofthesand,theflymaturesfully。Letussetheratliberty。ShewillgoandjointheothersontheSnakesinmypans。

CHAPTERXITHEBUMBLEBEEFLY

Underneaththewasp’sbrownpapermanorhouse,thegroundischanneledintoasortofdrainfortherefuseofthenest。Hereareshotthedeadorweaklylarvaewhichacontinualinspectionrootsoutfromthecellstomakeroomforfreshoccupants;here,atthetimeoftheautumnmassacre,areflungthebackwardgrubs;

here,lastly,liesagoodpartofthecrowdkilledbythefirsttouchofwinter。DuringtherackandruinofNovemberandDecember,thissewerbecomescrammedwithanimalmatter。

Suchricheswillnotremainunemployed。Theworld’sgreatlawwhichsaysthatnothingedibleshallbewastedprovidesfortheconsumptionofamereballofhairdisgorgedbytheowl。Howshallitbewiththevaststoresofaruinedwasps’nest!Iftheyhavenotcomeyet,theconsumerswhosetaskitistosalvethisabundantwreckagefornature’smarkets,theywillnottarryincomingandwaitingforthemannathatwillsoondescendfromabove。Thatpublicgranary,lavishlystockedbydeath,willbecomeabusyfactoryoffreshlife。Whoaretheguestssummonedtothebanquet?

Ifthewaspsflewaway,carryingthedeadorsicklygrubswiththem,anddroppedthemonthegroundroundabouttheirhome,thosebanqueterswouldbe,firstandforemost,theinsect—eatingbirds,thewarblers,allofwhomareloversofsmallgame。Inthisconnection,wewillallowourselvesabriefdigression。Weallknowwithwhatjealousintolerancethenightingalesoccupyeachhisowncantonment。Neighborlyintercourseamongthemistabooed。Themalesfrequentlyexchangedefiantcoupletsatadistance;but,shouldthechallengedpartydrawnear,thechallengermakeshimclearoff。Now,notfarfrommyhouse,inascantyclumpofhollyoakswhichwouldbarelygivethewoodcutterthewherewithalforadozenfaggots,Iused,allthroughthespring,tohearsuchfull—

throatedwarblingofnightingalesthatthesongsofthosevirtuosi,allgivingvoiceatonceandwithnoattemptatorder,degeneratedintoadeafeninghubbub。

Whydidthosepassionatedevoteesofsolitudecomeandsettleinsuchlargenumbersataspotwherecustomdecreesthatthereisjustroomenoughforonehouseholdonly?Whatreasonshavemadethereclusebecomeacongregation?Iaskedtheownerofthespinneyaboutthematter。

’It’slikethateveryyear,’hesaid。’TheclumpisoverrunbyNightingales。’

’Andthereason?’

’Thereasonisthatthereisahivecloseby,behindthatwall。’

Ilookedatthemaninamazement,unabletounderstandwhatconnectiontherecouldbebetweenahiveandthethrongingnightingales。

’Why,yes,’headded,’therearealotofnightingalesbecausetherearealotofbees。

Anotherquestioninglookfrommyside。Ididnotyetunderstand。

Theexplanationcame:’Thebees,’hesaid,’throwouttheirdeadgrubs。Thefrontofthehiveisstrewnwiththeminthemornings;

andthenightingalescomeandcollectthemforthemselvesandtheirfamilies。Theyareveryfondofthem。’

ThistimeIhadsolvedthepuzzle。Deliciousfood,abundantandfresheachday,hadbroughtthesongsterstogether。Contrarytotheirhabit,numbersofnightingalesarelivingonfriendlytermsinaclusterofbushes,inordertobenearthehiveandtohavealargershareinthemorningdistributionofplumpdainties。

Inthesameway,thenightingaleandhisgastronomicalrivalswouldhaunttheneighborhoodofthewasps’nests,ifthedeadgrubswerecastoutonthesurfaceofthesoil;butthesedelicaciesfallinsidetheburrowandnolittlebirdwoulddaretoenterthemurkycave,eveniftheentrancewerenottoosmalltoadmitit。Otherconsumersareneededhere,smallinsizeandgreatindaring;theflyiscalledforandhermaggot,thekingofthedeparted。Whatthegreenbottles,thebluebottlesandthefleshfliesdointheopenair,attheexpenseofeverykindofcorpse,otherflies,narrowingtheirprovince,doundergroundattheWasps’expense。

Letusturnourattention,inSeptember,tothewrapperofawasps’

nest。Ontheoutersurfaceandtherealone,thiswrapperisstrewnwithamultitudeofbig,white,ellipticaldots,firmlyfixedtothebrownpaperandmeasuringabouttwomillimetersandahalflongbyoneandahalfwide。Flatbelow,convexaboveandofalustrouswhite,thesedotsresembleveryneatdropsfallenfromatallowcandle。Lastly,theirbacksarestreakedwithfainttransversallines,anelegantdetailperceptibleonlywiththelens。Thesecuriousobjectsarescatteredalloverthesurfaceofthewrapper,sometimesatadistancefromoneanother,sometimesgatheredintomoreorlessdensegroups。TheyaretheeggsoftheVolucella,orbumblebeefly(Volucellazonaria,LIN。)

AlsostucktothebrownpaperoftheouterwrapperandmixedupwiththeVolucella’sarealargenumberofothereggs,chalkwhite,spear—shapedandridgedlengthwisewithsevenoreightthinribs,afterthemanneroftheseedsofcertainUmbelliferae。Thefinishingtouchtotheirdelicatebeautyisthefinestipplingalloverthesurface。Theyaresmallerbyhalfthantheothers。I

haveseengrubscomeoutofthemwhichmighteasilybetheearlieststageofsomepointedmaggotswhichIhavealreadynoticedintheburrows。Myattemptstorearthemfailed;andIamnotabletosaywhichflytheseeggsbelongto。Enoughforustonotethenamelessoneinpassing。Thereareplentyofothers,whichwemustmakeupourmindstoleaveunlabelled,inviewofthejumbledcrowdoffeastersintheruinedwasps’nest。Wewillconcernourselvesonlywiththemostremarkable,inthefrontrankofwhichstandsthebumblebeeFly。

Sheisagorgeousandpowerfulfly;andhercostume,withitsbrownandyellowbands,showsavagueresemblancetothatofthewasps。

OurfashionabletheoristshaveavailedthemselvesofthisbrownandyellowtocitetheVolucellaasastrikinginstanceofprotectivemimicry。Obliged,ifnotonherownbehalf,atleastonthatofherfamily,tointroduceherselfasaparasiteintothewasp’shome,sheresorts,theytellus,totrickeryandcraftilydonshervictim’slivery。Onceinsidethewasps’nest,sheistakenforoneoftheinhabitantsandattendsquietlytoherbusiness。

Thesimplicityofthewasp,dupedbyaveryclumsyimitationofhergarb,andthedepravityofthefly,concealingheridentityunderacounterfeitpresentment,exceedthelimitsofmycredulity。ThewaspisnotsosillynortheVolucellasocleverasweareassured。

IfthelatterreallymeanttodeceivetheWaspbyherappearance,wemustadmitthatherdisguiseisnonetoosuccessful。Yellowsashesroundtheabdomendonotmakeawasp。Itwouldneedmorethanthatand,aboveall,aslenderfigureandanimblecarriage;

andtheVolucellaisthicksetandcorpulentandsedateinhermovements。Neverwillthewasptakethatunwieldyinsectforoneofherownkind。Thedifferenceistoogreat。

PoorVolucella,mimesishasnottaughtyouenough。Youought——thisistheessentialpoint——tohaveadoptedawasp’sshape;andthatyouforgottodo:youremainedafatfly,easilyrecognizable。

Nevertheless,youpenetrateintotheterriblecavern;youareabletostaythereforalongtime,withoutdanger,astheeggsprofuselystrewnonthewrapperofthewasps’nestshow。Howdoyousetaboutit?

Letus,firstofall,rememberthatthebumblebeeflydoesnotentertheenclosureinwhichthecombsareheaped:shekeepstotheoutersurfaceofthepaperrampartandtherelayshereggs。Letus,ontheotherhand,recallthePolistes[atreenestingwasp]

placedinthecompanyofthewaspsinmyvivarium。Hereofasuretyisonewhoneednothaverecoursetomimicrytofindacceptance。Shebelongstotheguild,sheisawaspherself。Anyofusthathadnotthetrainedeyeoftheentomologistwouldconfusethetwospecies。Well,thisstranger,aslongasshedoesnotbecometooimportunate,isquitereadilytoleratedbythecagedwasps。Noneseekstopickaquarrelwithher。Sheisevenadmittedtothetable,thestripofpapersmearedwithhoney。Butsheisdoomedifsheinadvertentlysetsfootuponthecombs。Hercostume,hershape,hersize,whichtallyalmostexactlywiththecostume,shapeandsizeofthewasp,donotsaveherfromherfate。

SheisatoncerecognizedasastrangerandattackedandslaughteredwiththesamevigorasthelarvaeoftheHylotomasawflyandtheSaperdabeetle,neitherofwhichbearsanyoutwardresemblancetothelarvaofthewasps。

SeeingthatidentityofshapeandcostumedoesnotsavethePolistes,howwilltheVolucellafare,withherclumsyimitation?

Thewasp’seye,whichisabletodiscernthedissimilarinthelike,willrefusetobecaught。Themomentsheisrecognized,thestrangeriskilledonthespot。Astothatthereisnottheshadowofadoubt。

Intheabsenceofbumblebeefliesatthemomentofexperimenting,I

employanotherfly,Milesiafulminans,who,thankstoherslimfigureandherhandsomeyellowbands,presentsamuchmorestrikinglikenesstothewaspthandoesthefatVolucellazonaria。Despitethisresemblance,ifsherashlyventureonthecombs,sheisstabbedandslain。Heryellowsashes,herslenderabdomendeceivenobody。Thestrangerisrecognizedbehindthefeaturesofadouble。

Myexperimentsunderglass,whichvariedaccordingtothecaptureswhichIhappenedtomake,allleadmetothisconclusion:aslongasthereismorepropinquity,evenaroundthehoney,theotheroccupantsaretoleratedfairlywell;but,iftheytouchthecells,theyareassaultedandoftenkilled,withoutdistinctionofshapeorcostume。Thegrubs’dormitoryisthesanctumsanctorumwhichnooutsidermustenterunderpainofdeath。

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